Apteranthes (Caralluma) tuberculata
Herbarium sheet from the Royal Botanic Gardens, Kew¹
UPDATE: 16/9/2024
David Goyder of Kew Gardens (specialist in Angola, Madagascar and the Apocynaceae) has kindly looked into this for me. It seems the Kew Herbarium specimen is a combination of the specimens from Fleming and Stocks. The top row seem to be from Stocks. As the flowering specimen and "SEE SPIRIT COLLECTION" envelope are clearly from Stocks, the type specimen is Stocks 596. The catalogue should now reflect that Fleming's specimens are also on this sheet. I haven't altered the article below yet.
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The herbarium sheet above is the holotype specimen that was used to define the species
Caralluma tuberculata when that species was first described by the Kew Gardens botanist
Nicholas Edward Brown in 1892.² The plant was renamed
Apteranthes tuberculata in 2002 but this herbarium specimen still decides what plants are included in that species. If a plant is too different from this herbarium specimen it must be a different species.
When I saw this specimen I realised I had seen the handwriting before and it did not look like the handwriting or style of label used by the plant collector it was supposed to be. The collection was credited to Dr John Ellerton Stocks and given the collection number 596.¹
I believe it should be credited to Dr Andrew Fleming, a less well-known collector who also worked for the English East India Company in what is now Pakistan. He collected far fewer plant samples than Stocks but his main job at the time he collected this plant was a geological survey.
When referenced in botanical articles, a herbarium sheet is called by the name of the collector and the collection number. NE Brown referred to the Kew Herbarium specimen as Stocks 596, so it must have been misfiled at some time before 1892. This one should be called Fleming 78. I believe I have some convincing proof.